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Members Storage is a place are where members can store smaller items in 35 litre boxes. See Storage Hierarchy for more details on how to get your own box. In short: you can purchase a box through HMS and store it in here or downstairs members storage. The full rules about storage are available on the Member's Guide.
Upstairs storage has room for 129 boxes. It's mostly full at this point. There's plenty of space downstairs. There are some stats here.
See also
Large Project Storage - a place are where members can temporarily store projects that are being worked on
Members Storage Downstairs - another place where members will be able store smaller items in 35 litre boxes